For 35 years, Frankie, Linda, Kath, Brett, and Ally have met every Wednesday at the park near their homes in Palo Alto, California. Defined when they first meet by what their husbands do, the young homemakers and mothers are far removed from the Summer of Love that has enveloped most of the Bay Area in 1967. These "Wednesday Sisters" seem to have little in common: Frankie is a timid transplant from Chicago, brutally blunt Linda is a remarkable athlete, Kath is a Kentucky debutante, quiet Ally has a secret, and quirky, ultra-intelligent Brett wears little white gloves with her miniskirts. But they are bonded by a shared love of both literature - Fitzgerald, Eliot, Austen, du Maurier, Plath, and Dickens - and the Miss America Pageant, which they watch together every year.
As the years roll on and their children grow, the quintet forms a writers circle to express their hopes and dreams through poems, stories, and, eventually, books. Along the way, they experience history in the making: Vietnam, the race for the moon, and a women's movement that challenges everything they have ever thought about themselves, while at the same time supporting one another through changes in their personal lives brought on by infidelity, longing, illness, failure, and success.