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Only mutual communication can forge an alliance with the planet's sentient species and prove that mammals are more than tools. Forced to land on a planet they aren't prepared for, human colonists rely on their limited resources to survive. The planet provides a lush but inexplicable landscape—trees offer edible, addictive fruit one day and poison the next, while the ruins of an alien race are found entwined in the roots of a strange plant. Conflicts between generations arise as they struggle to understand one another and grapple with an unknowable alien intellect.

Over two hundred years after the first colonists landed on Pax, a new set of explorers arrives from Earth on what they claim is a temporary scientific mission. But the Earthlings misunderstand the nature of the Pax settlement and its real leader. Even as Stevland attempts to protect his human tools, a more insidious enemy than the Earthlings makes itself known. Stevland is not the apex species on Pax.