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When the Dearly's Dalmatians have their first litter of puppies (fifteen in all) everyone is delighted. But their joy is shattered when the pups are kidnapped! The humans don't have a clue as to who the culprit is, but the smartest dogs in London are on the case, and on the trail of fur-loving Cruella de Vil, who will stop at nothing to have a Dalmatian fur coat!

Pongo and Missis Pongo are a pair of Dalmatians who live with the newly married Mr. and Mrs. Dearly and their two nannies. Mr. Dearly is a "financial wizard" who has been granted life-long tax exemption and lent a house on the Outer Circle in Regent's Park in return for wiping out the government debt. The dogs consider the humans their pets, but allow the humans to think that they are the owners. Missis gives birth to a litter of 15 puppies.