Cat brothers Cecil and Anton are as different in personality as they are in looks. Cecil, stocky and black with white patches, thirsts for seafaring adventure as he roams the docks of his harborside home, taking day trips on fishing boats when the chance comes along. Slim, gray Anton prefers listening to the sailors' shanties at the town saloon. But one day when Anton does venture to port, he's stolen off the dock, and the next thing he knows, he's at sea.
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Terrible news has reached cat brothers Anton and Cecil: their rodent friend Hieronymus has been captured. Anton and Cecil must try to rescue the mouse who once saved Anton's life. Boarding one of the monstrous machines the mice call "landships," the brothers travel to the Wild West.
Tuckered out from a journey across the Wild West, cat brothers Anton and Cecil are ready to head east for home - until a minor stop to change trains in Chicago turns into a major escapade. The brothers are recruited by a police bloodhound to help solve a case: puppies are disappearing right off their leashes! Anton and Cecil's search takes them deep into the heart of the 1893 Chicago World's Fair, where they befriend exotic animals, ride the newly invented Ferris Wheel, and look for clues amid the crowds of fairgoers.